Knema oblongata Merr., J. Straits Branch Roy. Asiat. Soc. 85 (1922)

Synonyms
Knema obovoidea Merr.

Diagnostics
Sub-canopy tree up to 24 m tall and 46 cm dbh. Bark of trunk smooth, sometimes scaly or +/- fissured, once lenticellate; inner bark red-brown, beef-red or pinkish; slash wood pale yellowish, pale brownish, or white. Perianth lemon-yellow to red inside. Fruits yellow or orange, reddish hairy.

Description
Shrub or tree 4-20(-24) m. Twigs sometimes faintly angular, 2-5(-8) mm diameter, at first with rusty hairs 0.2-0.6 mm long, glabrescent; bark (finely) striate, not cracking or flaking. Leaves chartaceous or coriaceous, elliptic to (oblong-)lanceolate, 11-45 by 2.5-14 cm, base subattenuate, rounded, or shallowly cordate, apex acute(-acuminate); drying greenish brown, +/- glossy above, with scattered hair scars (lens!), lower surface greyish, finely papillate, with yellow-brown to rusty hairs of variable size, persistent or partially late glabrescent; dots absent; midrib raised above; nerves (10-)18-26 pairs, sunken or raised above; venation fine, distinct; petiole late glabrescent, 10-25 by 2-6 mm. Inflorescences: peduncle up to 4 mm, brachyblast simple or forked, to 5 mm long, in male 5-25-flowered, female 1-5-flowered; flowers with rusty hairs 0.1-1(- 2) mm long; perianth 3- (or 4-)lobed, yellow (?) or red inside. Male flowers: pedicel 2- 8 mm long, bracteole caducous, median to subapical; buds depressed broadly obovoid, tapering in the lower half, grooved when dry, 2.5-5 by 2.5-5 mm, cleft l/2-2/3(-3/4), lobes 0.5-1 mm thick; staminal disc circular or faintly triangular, flat or slightly concave, 1.5-3 mm diameter; anthers 7-9, or 13-18(-21), +/- half-sessile to just stiped, +/- oblique or suberect, (0.3-)0.5-0.7 mm. not or but slightly touching; androphore tapering or not, 1-2 mm long. Female flowers: pedicel 2.5-10 mm long, bracteole at or above the middle; buds ellipsoid to obovoid. 6-7.5 by 4-6 mm, cleft c. 1/2. lobes 0.5-1 mm thick; ovary subglobose, 3-3.5 mm diameter; style 1-1.5 mm; stigma 2-lobed and each lobe again 3-5- lobulate. Fruits 1-3 per infructescence, subovoid or obovoid, apex subacute to obtuse, base attenuate to rounded. 2-3.8 by 1.5-2 cm, with rusty hairs 1-1.5 mm long, easily rubbed off; dry pericarp 2-4 mm thick; fruiting pedicel 3-10 mm long. [from Flora Malesiana]

Ecology
In undisturbed to slightly disturbed (open) mixed dipterocarp, keranga and sub-montane forests up to 1200 m altitude. On hillsides and ridges with sandy soils.

Distribution
Borneo.

Local names
Borneo: Darah-darah, Kumpang.