Dysoxylum rugulosum King, J. As. Soc. Beng. 64, 2 (1895)

Latin for 'finely ribbed'.

Synonyms
Dysoxylum fulvum Airy Shaw
Dysoxylum undulatum Hend.

Diagnostics
Understorey tree up to 20 m tall and 39 cm dbh. Stipules absent. Leaves alternate, compound, leaflets penni-veined, glabrous. Flowers ca. 15 mm diameter, yellowish, placed in racemes. Fruits ca. 24 mm diameter, orange-red, fleshy, dehiscent capsule.

Description
Small tree to 20 m, flowering when a sapling; hole to 39 cm diam. Bark smooth, brown, to finely fissured and scaling; inner bark yellow-brown, mottled; sapwood cream. Branches striate, cicatrose. Leafy twigs 4-6 mm diam., fulvous tomentellous when young; apical hud stiletto-shaped. Leaves 20-40 cm, (1) 2-4 (5)-jugate, with a temiinal pair of leallets with a scar between them or one leallet and a spike or its scar; petiole 6-15 cm, +/- finely puberulous, flatened adaxially, conspicuously swollen at base; rachis angled or weakly winged. Leaflets 10-27 by 2.5-8 cm, the most distal the largest, oblong-lanceolate, chartaccous to subcoriaceous, rugulose, minutely black gland-dotted, alternate to sub-opposite, bases acute, attenuate, apices abruptly acuminate, costae 8-14 on each side, sometimes with domatia in axils, acute to obtuse, arcuate, prominent abaxially, secondary venation obscure; petiolules 4-10 mm, swollen, blackish in sicco. Thyrses 1-8 cm, spike-like, supra-axillary; rachis c. 2-3 mm diam., puberulous, bearing congested cymules of 3 or 4 sweetly scented flowers; bracts c. 1 mm, triangular. Calyx c. 2 mm long, 5 mm diam., very shallowly cupular, puberulous without, shortly 4-toothed. Petals 4, c. 8 by 3 mm, elliptic, puberulous without, yellowish, valvate. Staminal tube +/- 4-angled, glabrous or sparsely puberulous on angles without, margin crenate to (7-)8(-9)-toothed, the teeth emarginate; anthers (7) 8 (9), c. 1 mm long, elliptic, included. Disk c. 2 mm long and in diam., fleshy, glabrous or +/- pubescent within, margin crenate to c. 8-toothed. Ovary pubescent, 4-locular; style puberulent in proximal half; stylehead discoid with basal annulus. Capsule solitary or paired, 3-5 cm long. c. 2-2.5 cm diam., deeply 3- or 4-lobed, glabrous and orange when mature, drying blackish, veined. Seeds black. [from Flora Malesiana]

Ecology
In undisturbed mixed dipterocarp and sub-montane forests up to 2200 m altitude. On hillsides and ridges, but also on alluvial sites near or along rivers and streams. On sandy to ultramafic soils, but also on clay.

Distribution
Peninsular Malaysia, Sumatra, Borneo.

Local names
Borneo: Segarah, Segera.