Chisocheton pentandrus (Blanco) Merr., Philipp. Govt. Lab. Bur. Bull. 27 (1905)

Synonyms
Chisocheton beccarianus (Baill.) Harms
Chisocheton microcarpus Koord. & Valet.
Chisocheton microcarpus var. moluccanus Valet.
Chisocheton parvifolius Merr.
Chisocheton paucijugus (Miq.) B.D.Jackson
Chisocheton philippinus (Turcz.) Harms
Chisocheton sorsogonensis Elmer ex Merr.
Chisocheton spicatus Hiern
Dasycoleum beccarianum Baill.
Dasycoleum philippinum Turcz.
Schizochiton paucijugum Miq.
Trichilia pentandra Blanco

Diagnostics
Sub-canopy tree up to 26 m tall and 56 cm dbh. Stipules absent. Leaves alternate, compound, with leaf tip growing indeterminately, leaflets penni-veined, glabrous. Flowers ca. 10 mm diameter, yellowish, placed in panicles. Fruits ca. 17 mm diameter, red, capsules.

Description
Tree or treelet 3-18(-40) m tall and up to 56 cm diameter; buttressed to 60 cm. Bark greenish grey; inner bark pale fawn or pinkish; sapwood pale cream. Leafy twigs 2.5-6 mm diam., deciduously tawny pubescent to subglabrous. Leaves to 45 cm and to 9-jugate; petiole 2-10 cm, terete, minutely pubescent. Leaflets to 16.5(-26.5) by 6(-9) cm, elliptic to ovate-oblong, dark green adaxially, paler abaxially, glabrous or sparsely pubescent on veins, bases +/- asymmetric, acute to obtuse, apices acuminate to acutely cuspidate, costae 8-16 on each side; petiolules to 8 mm long. Inflorescence spiciform to thyrsoid, to 63 cm, axillary to supra-axillary, sometimes in axils of unexpanded leaves; axis finely velvety puberulous; flowers pedicellate, +/- fragrant. Calyx c. 4 mm long, +/- sparsely puberulous without, margin truncate to obscurely or irregularly lobed. Petals (4 or) 5, 8-12(-16) by 2 mm, cream, densely fulvescent-hirsute without, valvate, apex acute. Staminal tube white, +/- densely pilose, rarely subglabrous within, pubescent without; margin 5-lobed, lobes laciniate; anthers 5 (6), 3 mm long, glabrous. Ovary 2-locular, shortly stipitate, hirsute; style glabrous to pubescent. Infructescence to 30 cm. Capsule to 21 mm diam., globose or beaked, dull red, minutely rusty tomentose; pericarp with white latex. Seeds 2, to 15 mm diam., flattened, sarcotestal.

Ecology
In undisturbed mixed dipterocarp forests up to 400 m altitude. Usually on hillsides and ridges with sandy soils, but also on limestone.

Distribution
Peninsular Thailand, Peninsular Malaysia, Sumatra, Java, Lesser Sunda Islands, Borneo, Philippines, Celebes, Moluccas.

Local names
Borneo: Landur, Rusurang, Sibalia.